The expression 4 > 5 > 3 evaluates to false. This is because comparison operators are evaluated from left to right. First, 4 > 5 is evaluated to false, since 4 is not greater than 5. Then, false > 3 is evaluated. In this context, false is coerced to the number 0, so the expression becomes 0 > 3, which is false. This example highlights the importance of understanding operator precedence and associativity in JavaScript, as well as being careful with chained comparison operators.